Long ago the Dark Knight Sparda rebelled against the dark emperor and waged a one-man war to save humanity. Thousands of years later, a religious organization worships this renowned savior and has taken it upon themselves to rid the world of all evil. Residing in Fortuna, this organization is known as "Order of the Sword," and is made up of an elite group of "Holy Knights." One day a young knight named Nero witnesses Sparda's legendary son Dante slaughter his fellow warriors. Has the renowned devil hunter turned his back on mankind? What does this encounter mean for these two individuals?